A MOST PRODUCTIVE SMITHSONIAN-JCU COLLABORATION
Since 1998 I have travelled more-or-less annually to Panama to collaborate with Klaus Winter, a Senior Staff Scientist at the Smithsonian Tropical Research Institute (STRI).
Klaus and I have co-authored more than 30 scientific papers! We are collaborating on many of the experiments listed in the Experiments page.
Klaus Winter. Image from STRI website. 
Winterdom, Gamboa, Panama. Image from STRI website 
Erika Edwards, Joe Holtum, Andrew Smith, Howard Griffiths and Klaus Winter sampling Rowan Sage’s homebrew. Lake Tahoe. 
Jorge Aranda and Joe Holtum. Fortuna, Panama. 
Klaus Winter, Andrew Smith and a barrel cactus coping admirably with the Arizona Desert.
Joe Holtum's Science Site 